Miniature scarabs of the genus Haroldius on Sulawesi, with notes on their relatives (Coleoptera: Scarabaeidae)

Abstract

Haroldius Boucomont, an Oriental-Afrotropical genus of unusually small Scarabaeinae, is re-diagnosed, and recorded from the island of Sulawesi (Indonesia) for the first time. Five new species from North and Southeast Sulawesi are described, pictured, and keyed (Haroldius cambeforti, H. celebensis, H. kolaka, H. penelopae, H. tangkoko). Their characters are analyzed and compared with those of the Sundaland species. An annotated list of the species currently placed in Haroldius and some related genera is given. The Sundaland and Sulawesi species of Haroldius are all diagnosed. Outstanding problems with the systematic position of Haroldius and related genera are discussed, and their world distribution is mapped.