Original Description of the Brown Frog from Hokkaido, Japan (Genus Rana)

Abstract

Rana pirica, a brown frog from Hokkaido, Japan, is described as a new species. The species is closely related to R. ornativentris, R. dybowskii and R. chensinensis, characterized by 2n=24 chromosomes, but is distinct morphologically from all three, and genetically from the first two species. Systematic problems of brown frogs from east Asia are discussed.