The genital anatomy of Orcula jetschini (Romania), O. zilchi (Bulgaria) and O. wagneri (Albania) is described. Based on anatomical features (morphology of the penial caecum), shell characters (sculpture and shape) and unpublished molecular data, the genus Orcula is subdivided into three subgenera. O. zilchi was classified within the monotypic subgenus Orcula (Hausdorfia) subgen. n.; O. jetschini, O. wagneri and O. schmidtii were classified to Orcula (Illyriobanatica) subgen. n. (type species: Pupa Schmidtii), whereas the other Orcula species remain in the nominotypical subgenus. Orcula (Hausdorfia) is known from South-Eastern Bulgaria and North-Western Turkey, Orcula (Illyriobanatica) inhabits Western Romania, North-Western Greece, Albania, Macedonia, Kosovo and Montenegro. The nine species of Orcula (Orcula) are known mainly from the Alps and the Western Carpathians (from Eastern France to Eastern Hungary and Slovakia). The occurrence of only one Orcula species, namely O. jets chini is verified from Romania. Available information suggests that data on the Romanian occurrence of Orcula dolium and O. gularis were based on wrongly identified specimens. Sphyradium dobrogicum (=Orcula dobrogica) is considered as a synonym of S. doliolum. © Barna Páll-Gergely et al.
Páll-Gergely, B., Deli, T., Irikov, A., & Harl, J. (2013). Subgeneric division of the genus Orcula Held 1837 with remarks on Romanian orculid data (Gastropoda, Pulmonata, Orculidae). ZooKeys, 301, 25–49. https://doi.org/10.3897/zookeys.301.5304