Behavior strongly suggests 2-4 locus systems in grasses and Ranunculaceae, Chenopodaceae; discussion of whether rejection depends on match of any allele or all, efficiency argument for latter in terms of allowing cross-compatibility. Several reasons given why complex S-I may be widespread and easy to miss, e.g., few spp. studied genetically in detail, mind set for single locus S-I, crosses not wide enough to reveal underlying genetics, distinction of recognition and response loce (not called such but this is essentially what Denward proposed in 1963; see p. 241). Proposes an effectively oppositional model for action of multilocus S-I with all allele match needed for response, but mentions empirical problems with the model including pseudo S-I.
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