An unusual new species of narcomedusa from Tasmania is described. It differs from all other species of narcomedusae in having two whorls of tentacles, as well as a beveled opening in the apical surface containing peculiar 'sky-scraper-like' protruding structures, which we have interpreted as gonads. We place this unusual form in its own family, the Csiromedusidae fam. nov. Knowledge about its basic biology and ecology is currently lacking. We provide a comparative table of features of the families of Narcomedusae, as well as a summary of characters observed in known juvenile and larval stages of species of Narcomedusae.
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