Abstract. The homology of veins and other wing characters in Heteroptera is reviewed in the light of palaeontology and new functional studies. A cladogram is given for the higher taxa of Hemiptera. It is probable that the vannus is an autapomorphy of Auchenorrhyncha+Heteropteroidea; that the leading edge vein of heteropteran fore- and hindwings is C+Sc; that Rs cannot be distinguished from R; that the hamus is part of M; that the glochis is a secondary structure. The difficulty of defining a vein is stressed. The functional significance of the hemielytron, cuneal fracture and longitudinal flexion lines is discussed. A preliminary ground-plan for Heteroptera wings is presented.
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