On the basis of morphoanatomical parameters, the sand smelt species (Atherina boyeri Risso, 1810) is viewed as a highly polymorphic complex. In this study, intraspecific sequence variation in a portion of the cytochrome b gene was examined in 88 individuals from Tunisia and France. The correlation between the results of statistical analysis of the sequence data using a variety of tree-building algorithms and morphoanatomical analyses demonstrated the subdivision into three putative species: A. boyeri, which only includes non-punctuated fishes, A. punctata, which corresponds to punctuated fishes and A. lagunae, which corresponds to atherines living in lagoons. © 2002 Académie sciences / Éditions scientifiques et médicales Elsevier SAS.
Trabelsi, M., Gilles, A., Fleury, C., Mâamouri, F., Quignard, J. P., & Faure, É. (2002). Atherina punctata and Atherina lagunae (Pisces, Atherinidae), new species found in the Mediterranean Sea. 2. Molecular investigations of three Atherinid species. Comptes Rendus - Biologies, 325(11), 1119–1128. https://doi.org/10.1016/S1631-0691(02)01529-9