A countrywide survey in Oman revealed Middle East respiratory syndrome coronavirus (MERS-CoV) nucleic acid in five of 76 dromedary camels. Camel-derived MERS-CoV sequences (3,754 nucleotides assembled from partial sequences of the open reading frame (ORF)1a, spike, and ORF4b genes) from Oman and Qatar were slightly different from each other, but closely related to human MERS-CoV sequences from the same geographical areas, suggesting local zoonotic transmission. High viral loads in nasal and conjunctival swabs suggest possible transmission by the respiratory route.
Nowotny, N., & Kolodziejek, J. (2014). Middle East Respiratory Syndrome coronavirus (MERS-CoV) in dromedary camels, Oman, 2013. Eurosurveillance, 19(16). https://doi.org/10.2807/1560-7917.ES2014.19.16.20781