A resistant phenotype similar to that conferred in wheat by the complementary genes Lr27 + Lr31 was produced in the progeny of intercrosses of cultivars carrying Lr27 and a line possessing Lr12. This confirms that Lr12 is either completely linked with Lr31 or is the same gene. On the basis of these findings and that Lr31 is located on chromosome 4BS, it is concluded that Lr12 must also be located on 4BS. Adult-plant genetic tests confirm that the Australian wheat cultivar Timgalen carries Lr12, and stocks with Lr12 alone were established from this cultivar.
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Singh, D., Park, R. F., & McIntosh, R. A. (1999). Genetic relationship between the adult plant resistance gene Lr12 and the complementary gene Lr31 for seedling resistance to leaf rust in common wheat. Plant Pathology, 48(5), 567–573. https://doi.org/10.1046/j.1365-3059.1999.00391.x