A morphological review of subspecies of the Asian box turtle,Cuora amboinensis (Testudines, Geomydidae)

Citations of this article
Mendeley users who have this article in their library.
Get full text


The turtle Cuora amboinensis has an extensive distribution covering most of southern mainland Asia, Indonesia, and extending to the Philippine Islands. Unlike many species, C. amboinensis occurs on both sides of Wallace's Line separating Asian and Australian flora and fauna. Four subspecies are currently recognized; Cuora a. kamaroma (southern continental Asia, Java and the northern Philippines "introduced"), C. a. lineata (Kachin Province, Myanmar "Burma" and adjacent Yunnan Province, China), C. a. couro (Sumatra, Java, Sumbawa, and adjacent smaller Indonesian islands); and C. a. amboinensis (Moluccas, Sulawesi, Philippines). Five pattern and 33 morphological characters were examined for variation in 691 individuals from throughout the species' range. Our analyses suggest that only two presently recognized subspecies are valid: amboinensis and kamaroma. Neither couro nor lineata are supported by our analysis. We recommend that C. a. couro should be synonymized with the species C. amboinensis and C. a. lineata with the subspecies C. a. kamaroma.




Ernst, C. H., Laemmerzahl, A. F., & Lovich, J. E. (2016). A morphological review of subspecies of the Asian box turtle,Cuora amboinensis (Testudines, Geomydidae). Proceedings of the Biological Society of Washington, 129(1), 144–156. https://doi.org/10.2988/0006-324X-129.Q2.144

Register to see more suggestions

Mendeley helps you to discover research relevant for your work.

Already have an account?

Save time finding and organizing research with Mendeley

Sign up for free