Revision of the subfamily Thyreocorinae of the Pentatomidae (Hemiptera-Heteroptera)

Abstract

Thyreocorinae redefined, excluding Strombosoma and Carrabas, which are keyed (in appendix) but not given a new subfamily assignment; key to 9 genera: Galgupha, key to 15 subgenera; G. (Eury-scytus), key to 38 spp.; G. (E.) aliena (p.213), Brazil; G. (E.) apicata (p.213) British, French, and Dutch Guiana; G. (E.) assimilis* (p.214), Brazil; G. (E.) australis* (p.215), Brazil (type), Paraguay, Argentina, and Bolivia; G. (E.) basalis* (Germar) ( = Corime-laena proxima Walker); G. (E.) caudata [male], G. (E.) caudiculata [female], and G. (E.) contra [male] (p.217), Argentina; G. (E.) contracta [male] (p.218), Brazil; G. (E.) corvina (Euryscytus corvinus Horvath) (p.219); G. (E.) cru-enta (E. cruentus Horvath) (p.219); G. (E.) curvata [male] (p.220) and G. (E.) curvatula [female] (p.220), Brazil; G. (E.) difficilis* (Thyreocoris d. Breddin) (p.221); G. (E.) foveolata (Euryscytus foveolatus Jensen-Haarup) (p.222); G. (E.) insula (p.223), Paraguay and Argentina; G. (E.) labrata [female] (p.224), no locality; G. (E.) laevis* (Corimelaena l. Stal = C. integra Walker) (p. 224); G. (E.) lucretia* (p.225), Mexico (type) and Guatemala; G. (E.) mammula [female] (p.226), Brazil; G. (E.) nitida (p.226), Canal Zone (type) and Panama; G. (E.) obesa [male]* (p.227), Paraguay; G. (E.) opacifrons [male]* (p.227), Brazil; G. (E.) opercula [male]* (p.228), Mexico; G. (E.) parae (p.228), Brazil; G. (E.) parallela* (p.228), Brazil (type) and Bolivia; G. (E.) punctata [male]* (p.229), Mexico; G. (E.) rasilis (Euryscytus r. Horvath) (p.230); G. (E.) regularis [male]* (p.231), Argentina; G. (E.) romana (p.231), G. (E.) rostrata [male]* (p.232), and G. (E.) sahlbergi [male] (p.233), Brazil; G. (E.) signatipennis* (Odontoscelis s. Germar = Acrot-metus cetratus Horvath) (p.233); G. (E.) smidtii* (Tetyra s. Fab.) (p.234); G. (E.) valvata [female] (p.235), Guiana; G. (E.) varipennis [female]* (p.236), Brazil; G. (E.) vulnerata (Euryscytus v. Horv.) (p.236); BONARIA (p.237) subg. in Galgupha, type G. (B.) longirostris* (Thyreocoris l. Berg) (p.237); Galgupha (Gyrocnemis) key to 36 spp.; G. (Gyr.) acuta [female]* (p.246), Brazil; G. (Gyr.) basigrapha (Euryscytus basigraphus Horv.) (p.246); G. (Gyr.) bicornis [male]* (p.247), Brazil; G. (Gyr.) cruralis* (Corimelaena c. Stal = C. invaria Walk., Thyreocoris circumfusus Berg, T. borellii Montandon) (p.249); G. (Gyr.) cydnoidea ? (p.251), Colombia; G. (Gyr.) diminuta* (Euryscytus diminutus V. Duz.) (p. 252); G. (Gyr.) dimorpha* (p.253), Paraguay; G. (Gyr.) diversa [male] (p.254), Argentina and Paraguay; G. (Gyr.) geayi [female] (p.256), Venezuela; G. (Gyr.) gut-tiger* (Thyreocoris g. Stal); G. (Gyr.) impressa* (E. i. Horv.) (p.257); G. (Gyr.) inaequalis* (p.258), Colombia (type), Venezuela, and Bolivia; G. (Gyr.) intermedia [male] (p.258), Brazil; G. (Gyr.) lineata [male]* (p.259), Argentina (type) and Brazil; G. (Gyr.) maculipennis* (Odontoscelis m. Germar); G. (Gyr.) morbiloci [male] (p. 260), Mexico; G. (Gyr.) nitens [male]* (Thyreocoris n. Breddin) (p.260); G. (Gyr.) nitidipennis [female] (p.261), Venezuela; G. (Gyr.) notha and G. (Gyr.) novatra [female] (p.262), Argentina; G. (Gyr.) occulta (p.263), and G. (Gyr.) omissa [female] (p.263), Brazil; G. (Gyr.) parva* (p.264), Brazil and Paraguay; G. (Gyr.) punctifer (p. 264), Guatemala (type) Mexico, Canal Zone, Panama, Texas, Costa Rica, Brit. Honduras; G. (Gyr.) quadri-signata (Thyreocoris quadrisignatus Stal) (p.266); G. (Gyr.) reinhardti* (Euryscytus r. Jens.-Haarup) (p. 267); G. (Gyr.) singularis [female]* (p.267), Argentina; G. (Gyr.) soror [female] (p.268), Mexico; G. (Gyr.) tabida [female] (Corimelaena t. Stal) (p.268); G. (Gyr.) texana [male] (p.269), Texas; G. (Gyr.) triconcava* (p.269), Paraguay, Brazil. Bolivia, Argentina; G. (Gyr.) unica [female] (p.270), Colombia; CTENOPODA (p.270), subg. in Galgupha, type G. (C.) castor*, also includes G. (C.) pollux* (p.271), both Brazil; TREPOCNEMIS (p.272), subg. in Galgupha, type G. (T.) anomala [female]* (p.272), Argentina; OROCORIS (p.272), subg. in Galgupha, type G. (O.) arizonensis* (Cydnoides a. V. Duzee); G. (Acrotmetus), key to 6 spp.; G. (A.) brasilianus (A. b. Jensen-Haarup) (p.275); G. (A.) crassa (p.275), Brazil; G. (A.) schulzii* (Cimex s. Fabr. = Thyreocoris setiger Berg, Corimelaena major Breddin, Acrotmetus sphaeri-dioides Horv.) (p.277); G. (A.) tucumanus* (A. t. Horv. = A. persimilis Horv.) (p.278); subg. Galgupha, key to 7 spp.; G. (G.) carinata* (p.282), Louisiana (type), Texas, and Oklahoma to Virginia and Tennessee; G. (G.) denudata* (Corimelaena d. Uhler) (p. 284); G. (G.) hesperia (p.284), California; G. (Micro-compsus) daldorfii v. fabricii* (p.287), Brazil; G. (M.) vinculata* (Odontoscelis vinculatus Germar) (p.288), and its vars. insularis, Trinidad (type) and Venezuela, pictula (Corimelaena p. Walker), scymnoides* (G. s. Jensen-Haarup) (p.289), and surinamensis (p.290) Dutch Guiana; subg. Nothocoris, key to 26 spp.; G. (N.) akarna (p.295), Argentina; G. (N.) alutacea* (p.295), Brazil, G. (N.) anomala (p.296), Colorado; G. (N.) boliviana* (p.296), Bolivia; G. (N.) brevis [female] (p.297), Brazil; G. (N.) brunnipennis (Odontoscelis b. Germar) (p.298); G. (N.) caracasana (p.299), Venezuela; G. (N.) chilocoroides* (Corimelaena c. Walker) (p.299); G. (N.) eas [female] (p.301), Texas; G. (N.) fos-sula [female] (p.302), Mexico; G. (N.) grenadensis [female] (p.302), Colombia; G. (N.) inornata [female] (p.302), Argentina; G. (N.) juno [female] (p.303), Argentina; G. (N.) marginalis [male] (p.303), Colombia; G. (N.) marginicollis* Horv. ([long dash] G. meinerti Jensen-Haarup); G. (N.) microph-thalma [male] (p.305), Brazil; G. (N.) nitiduloides (Cimex n. Wolff = Corimelaena cyanea Uhler, C. cyaneonigra Walker, Galgupha violacea Jensen-Haarup) (p.305), key to 3 sspp.; G. nitiduloides ssp. caerulescens* (Thyreo-coris c. Stal) (p.307); G. n. ssp. texensis (p.308), Texas; G. (Nothocoris) oculata (p.309), Paraguay; G. parvula is questionably assigned to subg. Nothocoris; G. (N.) repetita [male] (p.311), G. (N.) stramineipennis [male] (p.312), and G. (N.) surda [female] (p.313), Brazil; subg. Psestophleps, key to 17 spp.; G. (P.) altera [female] (p.317), Brazil; G. (P.) bergiana* (p.318), Brazil (type), Paraguay, Argentina, and Peru; G. (P.) bisignata* (p.319), G. (P.) carbonata [female] (p.319), and G. (P.) casta [female] (p. 3.20), Argentina; G. (P.) costomaculata (p.320), Paraguay; G. (P.) fimbriata (p.321), Dutch Guiana; G. (P.) imitans [female] and G. (P.) inops (p.322), Brazil; G. (P.) mayana [male] (p.323), Honduras; G. (P.) media* (p.323), British and Dutch Guiana; G. (P.) mexicana (p.324), Mexico; G. (P.) obovata* (p.326), Argentina; G. (P.) pallipennis [female] (p.326), Brazil; G. (P.) porcata* Horv. ( = G. lugubris Horv.); G. (P.) tabellula (p.327), Dutch Guiana; PTERONOMOS (p.328), subg. in Galtupha, type G. (P.) oblonga* (p.331), Brazil (type), Paraguay, and Argentina; G. (P.) amitta [female] (p.329), Argentina; G. (P.) australis* (p.330), Paraguay and Argentina; G. (P.) china* (p.330), Peru (type), Bolivia, Argentina, and Brazil; G. (P.) punctifrons [male]* (p.332) and G. (P.) vittifrons [male]* (p.333), Argentina; CHARODA (p.33), subg. in Galgupha, type G. (C.) simplex* (p.333), Chile (type) and Brazil; G. (Astiro-dcrma) albipennis* ssp. ampliata (p.337), Brazil, Uruguay, and Argentina; G. (A.) breddini [male]* (p.337), Brazil; Cydnoides (Cydnoides) ciliatus ssp. orientis (p. 340), Florida (type), Texas, Minnesota, Kansas, Missouri, Nebraska, and Colorado; C. (C.) confusus [female] (p.341), New Mexico (type), Arizona, Texas, Mexico; SAYOCORIS (p.342), subg. in Cydnoides, type C. (S.) albipennis* (Thyreocoris a. Say= Corimelaena sayi V.D.) (p.343); C. (S.) peregrinus [female] (p.344). Lower California; COSMARIOIDES (p.345), subg. in Cydnoides, type C. (Cosm.) nitens [female] (p.345), Brazil, and also includes C. (Cosm.) reticulatus (p.346), Brazil and C. (Cosm.) setiventris* (p.346), Paraguay (type) and Bolivia; Amyssonotum rastratum* ssp. flexum* (p.351), Panama (type), Colombia, Canal Zone, Costa Rica, Guatemala, Mexico, and Texas; Pericrepis afer (p.352), Argentina; PRUHLERIA (p.354), type P. incerta [female]* (Corimelaena i. Uhler) (p.354); ALLOCORIS (p.358), type A. (A.) gillettii* (Corimelaena g. V.D.) (p.369); subgenus Allocoris, key to 22 spp.; A. (A.) agrella* (Corimelaena a. McAtee) (p.362); A. (A.) alpina* (p. 363), New York; A. (A.) alticola (Eucoria a. Horv.) (p.364); A. (A.) barberi* (p.365), Texas (type), Mexico, and Costa Rica; A. (A.) championi (Thyreocoris c. Distant) (p.365); A. (A.) contrasta [male] (p.366), Arizona; A. (A.) corallina (p.366), Guatemala (type), and Mexico; A. (A.) digitata [male] (p.367), Mexico; A. (A.) ele-gans* (p.368), Panama (type), Trinidad, Guatemala, Costa Rica, Mexico; A. (A.) feminea [female] (p.368), Texas; A. (A.) gillettii ssp. mexicana (p.370), Mexico; A. (A.) harti* (Corimelaena h. Malloch) (p.371); A. (A.) in-terrupta* (C. i. Mall.) (p.372); A. (A.) limata [female]* (p.373), no locality; A. (A.) marginella* (C. m. Dallas) (p.373); A. (A.) micans [male] (p.374), Guatemala; A. (A.) minuta (C. to. Uhler) (p.375); A. (A.) nigra* (C. n. Dallas = C. anthracina Uhler) (p.376); A. (A.) pal-meri* (p.377), Mexico; A. (A.) polita (C. p. Mall.); A. (A.) pulicaria* (Odontoscelis p. Germar = Galgupha flavo-marginata Thomas) (p.378); A. (A.) tibialis* (Tetyra t. Fabr.) (p.379); TERMAPORA (p.381), subg. in Allocoris, type A. (T.) minutissima* (Corimelaena m. Mall.); PARAPORA (p.382), subg. in Allocoris, type A. (P.) extensa* (C. e. Uhler= Thyreocoris montanus V.D.) (p.385); A. (P.) californica [female]* (C. c. V.D.) (p.383); A. (P.) cognata* (Thyreocoris cognatus V.D. = Eucoria placata Horv.) (p.384); A. (P.) incognita* (p.386), Arizona (type), from Colorado and British Columbia to Texas, California and Mexico; A. (P.) parana* (C. p. McAtee & Mall.) (p.387); A. (P.) virilis [male]* (p.387), Arizona (type), Washington, California, Idaho,