The basic ideas of genetic load and the cost of natural selection are both from J. B. S. Haldane. In his early papers on natural selection (1924--1932), Haldane was concerned with both the dynamics and the statics of evolution. He emphasized that, although evolution depends on changes of gene frequency, nevertheless at any one time the population is in approximate equilibrium for most factors.
CITATION STYLE
Crow, J. F. (1970). Genetic Loads and the Cost of Natural Selection. In Mathematical Topics in Population Genetics (pp. 128–177). Springer Berlin Heidelberg. https://doi.org/10.1007/978-3-642-46244-3_5
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