We have identified 6 Escherichia coli genomic genes, including 4 new genes, responsive to the stationary phase. One of them was regulated positively by RpoS at the stationary phase, and the remaining 5 negatively at a late stationary phase, all of them responding to multiple environmental stresses. Nucleotide sequences as well as such multiple responses revealed that those genes may have more than one overlapping-promoter recognized by different σ-factors which regulate gene expressions during their cell growth.
Talukder, A. A., Yanai, S., Nitta, T., Kato, A., & Yamada, M. (1996). RpoS-dependent regulation of genes expressed at late stationary phase in Escherichia coli. FEBS Letters, 386(2–3), 177–180. https://doi.org/10.1016/0014-5793(96)00426-7