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ZooKeys (, ISSN: 1313-2989 (print), ISSN: 1313-2970 (online)) is a peer-reviewed, open-access, online and print, journal launched to support free exchange of ideas and information in systematic zoology. All papers can be freely copied, printed and distributed at no charge. Authors are thus encouraged to post the pdf files of published papers on their homepages or elsewhere to expedite distribution. There is no charge for color. Published by Pensoft (

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