Two new genera of caeninae (Ephemeroptera: Caenidae), with a cladistic analysis of the tribe clypeocaenini

Abstract

The new genera Mandelara n. gen. (type species M. immutata n. sp.) from South Africa and Papuaenis n. gen. (type species P. balkei n. sp.) from Papua New Guinea are described. Both genera are included in the tribe Cly-peocaenini that is defined by a bulged frons and more or less protruding clypeus. A cladistic analysis recovers the Clypeocaeninae as monophyletic. Caenini + Tasmanocoenini are revealed as sister group to Clypeocanini, together constituting the subfamily Caeninae. A larval key in provided to the genera of Clypeocaenini.