Revision of the Agathidinae (Hymenoptera, Braconidae) of Vietnam, with the description of forty-two new species and three new genera

Abstract

The species of seventeen genera of Agathidinae (Braconidae) from Vietnam are revised: Agathis Latreille, 1804, Bassus Fabricius, 1804; Biroia Szépligeti, 1900; Braunsia Kriechbaumer, 1894; Camptothlipsis Enderlein, 1920; Coccygidium de Saussure, 1892; Coronagathis gen. n. (type species: C. cornifera sp. n.); Cremnops Foerster, 1862; Disophrys Foerster, 1862; Earinus Wesmael, 1837; Euagathis Szépligeti, 1900; Gyragathis gen. n. (type species: G. quyi sp. n.), Gyrochus Enderlein, 1920; Lytopylus Foerster, 1862; Therophilus Wesmael, 1837; Troticus Brullé, 1846, and Zelodia gen. n. (type species: Zelomorpha varipes van Achterberg & Maetô, 1990). Keys to the Vietnamese species are given. Sixty-five species are recognised, of which twelve species are newly recorded for Vietnam: Bassus albifasciatus (Watanabe, 1934), Coccygidium angostura (Bhat & Gupta, 1977), Cremnops atricornis (Smith, 1874), stat. n., Disophrys erythrocephala Cameron, 1900, Gyrochus yunnanensis Wang, 1984, Lytopylus romani (Shestakov, 1940), comb. n., Therophilus festivus (Muesebeck, 1953), comb. n., T. javanus (Bhat & Gupta, 1977), comb. n., T. lienhuachihensis (Chou & Sharkey, 1989), comb. n., T. marshi (Bhat & Gupta, 1977), comb. n., Zelodia absoluta (Chen & Yang, 1998), comb. n. and Z. longidorsata (Bhat & Gupta, 1977), comb. n. Forty-two species are new to science: Agathis citrinisoma sp. n., Bassus albobasalis sp. n., B. albozonatus sp. n., Biroia soror sp. n., Braunsia bicolorata sp. n., B. devriesi sp. n., B. maculifera sp. n., B. nigrapiculata sp. n., B. pumatica sp. n., Camptothlipsis hanoiensis sp. n., Coronagathis cornifera sp. n., Earinus aurantius sp. n., E. brevistigmus sp. n., Euagathis flavosoma sp. n., Disophrys maculifera sp. n., D. quymanhi sp. n., D. rhinoides sp. n., Gyragathis quyi sp. n., Therophilus annuliferus sp. n., T. cattienensis sp. n., T. contrastus sp. n., T. crenulisulcatus sp. n., T. depressiferus sp. n., T. elongator sp. n., T. levisoma sp. n., T. marucae sp. n., T. mellisoma sp. n., T. nigrolineatus sp. n., T. nuichuaensis sp. n., T. parasper sp. n., T. planifrons sp. n., T. punctiscutum sp. n., T. robustus sp. n., T. rugosiferus sp. n., T. scutellatus sp. n., Troticus alloflavus sp. n., T. giganteus sp. n., Zelodia albobasalis sp. n., Z. anginota sp. n., Z. bicoloristigma sp. n., Z. brevifemoralis sp. n. and Z. flavistigma sp. n. The following new synonyms are proposed: Euagathis nigrithorax Bhat & Gupta, 1977, E. variabilis Enderlein, 1920, E. variabilis var. tibialis Enderlein, 1920, E. variabilis var. melanopleura Enderlein, 1920 and E. variabilis var. sucarandana Enderlein, 1920 wiThe. abbotti (Ashmead, 1900); E. jinshanensis Chen & Yang, 2006 and E. sharkeyi Chen & Yang, 2006, wiThe. forticarinata (Cameron, 1899). The genus Amputostypos Sharkey, 2009, is synonymised with Coccygidium de Saussure, 1892, syn. n. The following new combinations are given: Bassus subrasa (Enderlein, 1920), comb. n., Gyragathis angulosa (Bhat & Gupta, 1977), comb. n., Lytopylus romani (Shestakov, 1940), comb. n., Therophilus annulus (Chou & Sharkey, 1989), comb. n., T. asper (Chou & Sharkey, 1989), comb. n., T. cingulipes (Nees, 1812), comb. n., T. daanyuanensis (Chen & Yang, 2006), comb. n., T. fujianicus (Chen & Yang, 2006), comb. n., T. javanus (Bhat & Gupta, 1977), comb. n., T. lanyuensis (Chou & Sharkey, 1989), comb. n., T. luzonicus (Bhat & Gupta, 1977), comb. n., T. muesebecki (Bhat & Gupta, 1977), comb. n., T. rudimentarius (Enderlein, 1920), comb. n., T. similis (Bhat & Gupta, 1977), comb. n., T. sungkangensis (Chou & Sharkey, 1989), comb. n., T. tanycoleosus (Chen & Yang, 2006), comb. n., T. tonghuaensis (Chen & Yang, 2006), comb. n., T. tongmuensis (Chen & Yang, 2006), comb. n., T. transcasperatus (Chen & Yang, 2006), comb. n., Troticus latiabdominalis (Bhat, 1978), comb. n., Zelodia absoluta (Chen & Yang, 1998), comb. n., Z. achterbergi (Chen & Yang, 2006), comb. n., Z. albopilosella (Cameron, 1908), comb. n., Z. chromoptera (Roman, 1913), comb. n., Z. nihonensis (Sharkey, 1996), comb. n., Z. cordata (Bhat & Gupta, 1977), comb. n., Z. diluta (Turner, 1918), comb. n., Z. dravida (Bhat & Gupta, 1977), comb. n., Z. exornata (Turner, 1918), comb. n., Z. longidorsata (Bhat & Gupta, 1977), comb. n., Z. longiptera (Yang & Chen, 2006), comb. n., Z. maculipes (Cameron, 1911), comb. n., Z. nigra (Bhat & Gupta, 1977), comb. n., Z. philippinensis (Bhat & Gupta, 1977), comb. n., Z. reticulosa (Yang & Chen, 2006), comb. n., Z. quadrifossulata (Enderlein, 1920), comb. n., Z. ruida (Sharkey, 1996), comb. n., Z. similis (Bhat & Gupta, 1977), comb. n., Z. penetrans (Smith, 1860), comb. n. and Z. varipes (van Achterberg & Maetô, 1990), comb. n. © C. van Achterberg, K.D. Long.